Contact Information

Chesapeake Biological Laboratory
P.O. Box 38
Solomons, MD 20688
Phone: 410-326-7266
Fax: 410-326-7318




Palinkas, C.M., J.M. Testa, J.C. Cornwell, M. li, and L.P. Sanford. 2019. Influences of a river dam on delivery and fate of sediments and particulate nutrients to the adjacent estuary: case study of Conowingo Dam and Chesapeake Bay. Estuaries and Coasts 42: 2072-2095.

Orth, R.J., W. Dennison, C. Gurbisz, M. Hannam, J. Keisman, J.B. Landry, J. Lefcheck, K. Moore, R. Murphy, C. Patrick, J.M. Testa, D. Weller, D. Wilcox, and R. Batiuk. 2019. Long-term annual aerial surveys of submersed aquatic vegetation (SAV) support science, management, and restoration. Estuaries and Coasts,

Shen, C., J.M. Testa, W. Ni, W.-J. Cai, M. Li, and W.M. Kemp. 2019. Ecosystem metabolism and carbon balance in Chesapeake Bay: A 30-year analysis using a coupled hydrodynamic-biogeochemical model. Journal of Geophysical Research - Oceans,

Liang, D., L.A. Harris, J.M. Testa, V. Lyubchich, and S. Filoso. 2019. Detection of the effects of stormwater control measure in streams using a Bayesian BACI power analysis. Science of The Total Environment 661: 386-392.

Shen, C., Testa, J.M., Li, M., Cai, W.‐J., Waldbusser, G. G., Ni, W., Cornwell, J., Brodeur, J., and Su, J. 2019. Controls on carbonate system dynamics in a coastal plain estuary: A modeling study. Journal of Geophysical Research: Biogeosciences, 124.

Brodeur, J.R., B. Chen, J. Su, Y.-Y. Xu, N. Hussain, K.M. Scaboo, Y.J. Zhang, J.M. Testa, and W.-J. Cai. 2019. Chesapeake Bay Inorganic Carbon: Spatial Distribution and Seasonal Variability. Frontiers in Marine Science 6.

Testa, J.M., V. Lyubchich, and Q. Zhang. 2019. Patterns and trends in Secchi disk depth over three decades in the Chesapeake Bay estuarine complex. Estuaries and Coasts 42(4): 927-943.

Saba G., K. Goldsmith, S. Cooley, D. Grosse, S. Meseck, W. Miller, B. Phelan, M. Poach, R. Rheault, K. St.Laurent, J.M. Testa, J. Weis, R. Zimmerman. 2019. Recommended Priorities for Research on Ecological Impacts of Ocean and Coastal Acidification in the U.S. Mid-Atlantic. Estuarine, Coastal and Shelf Science.

Lyubchich, V., T. Lebedeva, Tatiana, and J.M. Testa. 2019. A Data-driven Approach to Detecting Change Points in Linear Regression Models. Environmetrics


Lefcheck, J.S., R.J. Orth, W.C. Dennison, D.J. Wilcox, R.R. Murphy, J. Keisman, C. Gurbisz, M. Hannam, J.B. Landry, K.A. Moore, C.J. Patrick, J.M. Testa, D.E. Weller, R.A. Batuik. 2017. Nutrient reductions lead to unprecedented recovery of a temperate coastal ecosystem. Proceedings of the National Academy of Science, USA. doi: 10.1073/pnas.1715798115. *Awarded PNAS Cozzarelli Prize for excellence and originality

Testa, J.M., W.M. Kemp, and W.R. Boynton. 2018. Season-specific trends and linkages of nitrogen and oxygen cycles in Chesapeake Bay. Limnology and Oceanography: doi: 10.1002/lno.10823.

Testa, J.M., R.R. Murphy, D.C. Brady, and W.M. Kemp. 2018. Nutrient- and climate-induced shifts in the phenology of linked biogeochemical cycles in a temperate estuary. Frontiers in Marine Science:

Fennel, K., and  J.M. Testa. 2018. Biogeochemical controls on coastal hypoxia. Annual Review of Marine Science 11:105-130.


Testa, J.M., J.B. Clark, W.C. Dennison, E.C. Donovan, A.W. Fisher, W. Ni, M. Parker, D. Scavia, S.E. Spitzer, A.M. Waldrop, V.M.D. Vargas, G. Ziegler. 2017. Ecological forecasting and the science of hypoxia in Chesapeake Bay BioScience 61(7): 614-626.

Boynton, W.R., M.A.C. Ceballos Fraser, E.M. Bailey, C.L. Hodgkins, J.L. Humphrey, J.M. Testa. 2017. Nutrient and oxygen exchanges at the sediment-water interface: a global synthesis and critique of estuarine and coastal data. Estuaries and Coasts 41(2): 301-333.

Cai, W.-J., Huang, W.-J., Luther, G.W., Pierrot, D., Li, M., Testa, J.M., Xue, M., Joesoef, A., Mann, R., Brodeur, J., Xu, Y.-Y., Chen, B., Hussain, N., Waldbusser, G.G., Cornwell, J.C., Kemp, W.M. 2017. Redox reactions and weak buffering capacity lead to acidification in the Chesapeake Bay, Nature Communications 8(1).


Schaeffer, E. D., Testa, J. M, Gel, Y. R., and Lyubchich, V. 2016. A Data-Driven Approach to Dynamic Spatio-Temporal Clustering. Proceedings of the 6th International Workshop on Climate Informatics, Boulder, CO

Testa, J.M., E.E. Adams, E.W. North, and R. He. 2016. Modeling the influence of deep water application of dispersants on the surface expression of oil: a sensitivity study. Journal of Geophysical Research: Oceans. doi: 10.1002/2015JC011571.

Staehr, P.A., J.M. Testa, and J. Carstensen. 2016. Decadal changes in water quality and net productivity of a shallow Danish estuary following significant nutrient reductions. Estuaries and Coasts doi 10.1007/s12237-016-0117-x.

Testa, J.M., W.M. Kemp, L.A. Harris, R.J. Woodland, and W.R. Boynton. 2016. Challenges and directions for the advancement of estuarine ecosystem science. Ecosystems doi: 10.1007/s10021-016-0004-0.

Li, M., Y.J. Lee, J.M. Testa, Y. Li, W. Ni, W.M. Kemp, and D.M. Di Toro. 2016. What drives interannual variability of estuarine hypoxia: Climate forcing versus nutrient loading? Geophysical Research Letters 43(5) 2127-2134 doi 10.1002/2015GL067334.

Irby, I.D., M.A.M. Friedrichs, C.T. Friedrichs, A.J. Bever, R.R. Hood, L.W.J. Lanerolle, M.E. Scully, K. Sellner, J. Shen, J.M. Testa, M. Li, H. Wang, P. Wang, L. Linker, and M. Xia. 2016. Challenges associated with modeling low-oxygen waters in Chesapeake Bay a multiple model comparison. Biogeosciences 13, 2011-2028.


Testa J.M., D.C. Brady, J.C. Cornwell, M.S. Owens, L.P. Sanford, C.R. Newell C., S.E. Suttles, and R.I.E. Newell. 2015.Modeling the impact of floating oyster aquaculture on sediment-water nutrient and oxygen fluxes. Aquaculture Environment Interactions. 7: 205-222 doi: 10.3354/aei00151.

Ganju, N.K., M.J. Brush, B. Rashleigh, A.L. Aretxabaleta, P. del Barrio, J.S. Grear, L.A. Harris, S.J. Lake, G. McCardell, J. O’Donnell, D.K. Ralston, R.P. Signell, J.M. Testa, J.M.P. Vaudrey. 2015. Progress and challenges in coupled hydrodynamic-ecological estuarine modeling. Estuaries and Coasts. 39: 311-332 doi: 10.1007/s12237-015-0011-y.

Harris, L.A., C.L.S. Hodgkins, M.C.Day, D.Austin, J.M. Testa,W.Boynton, L. Van Der Tak, and N.W. Chen. 2015. Optimizing recovery of eurtropic estuaries: Impact of destratification and re-aeration on nutrient and dissolved oxygen dynamics. Ecological Engineering. 75:470-483. doi: 10.1016/j.ecoleng.2014.11.028


Testa, J.M., Y. Li, Y.-J. Lee, M. Li, D.C. Brady, D.M. Di Toro, W.M. Kemp, and J.J. Fitzpatrick. 2014. Quantifying the effects of nutrient Loading on dissolved O2 cycling and hypoxia in Chesapeake Bay using a coupled hydrodynamic-biogeochemical model. Journal of Marine Systems. 139: 139-158 doi:10.1016/j.jmarsys.2014.05.018.

Testa, J.M., and W.M. Kemp. 2014. Spatial and temporal patterns in winter-spring oxygen depletion in Chesapeake Bay bottom waters. Estuaries and Coasts. 37(6): 1432-1448 doi: 10.1007/s12237-014-9775-8.


Testa, J.M., D.C. Brady, D.M. Di Toro, W.R. Boynton, and W.M. Kemp. 2013. Sediment flux modeling: Nitrogen, phosphorus and silica cycles. Estuarine, Coastal and Shelf Science, 131:245-263.

Brady, D.C., J.M. Testa, D.M. Di Toro, W.R. Boynton, and W.M. Kemp. 2013. Sediment flux modeling: Calibration and application for coastal systems. Estuarine, Coastal and Shelf Science 117: 107-124.


Testa, J.M., W.M. Kemp, C.S. Hopkinson, and S.V. Smith. In Press. Ecosystem Metabolism. In: Day, J.W., Crump, B.C., Kemp, W.M., and Yáñez-Arancibia,A. (Eds.), Estuarine Ecology, Vol 2, Chapter 15. John Wiley & Sons, Inc., Hoboken, NJ.

Testa, J.M. and W.M. Kemp. 2012. Hypoxia-induced shifts in nitrogen and phosphorus cycling in Chesapeake Bay. Limnology and Oceanography 57: 835-850.

Stæhr, P.A., J.M. Testa, W.M. Kemp, J.J. Cole, K. Sand-Jensen, S.V. Smith. 2012. The metabolism of aquatic ecosystems: History, methods, and applications. Aquatic Sciences 74: 15-29.


Kemp, W.M. and J.M. Testa. 2011. Metabolic Balance between Ecosystem Production and Consumption. In: Wolansky, E. and McLusky, D.S. (eds.), Treatise on Estuarine and Coastal Science, 7: 83-118. Waltham: Academic Press.

Testa, J.M. and W.M. Kemp, 2011. Oxygen - Dynamics and Biogeochemical Consequences. In: Wolansky, E. and McLusky, D.S. (eds.), Treatise on Estuarine and Coastal Science, 5:163-199. Waltham: Academic Press.


Lookingbill, T., T.J.B. Carruthers, J.M. Testa, W.K. Nuttle, and G. Shenk. 2010. Environmental models: Providing synthesis, analysis, simulation, and prediction. In: B.J. Longstaff, T.J.B. Carruthers, W.C. Dennison, T.R. Lookingbill, J.M. Hawkey, J.E. Thomas, E.C. Wicks, and J. Woerner (eds.) Integrating and applying science: A handbook for effective coastal ecosystem assessment. IAN Press, Cambridge, MD, 133-149.

Testa, J.M., C. Gurbisz, L. Murray, W. Gray, J. Bosch, C. Burrell, and W.M. Kemp. 2010. Investigating Aquatic Dead Zones: A series of activities designed to explore a mystery of the deep. The Science Teacher 77(2): 29-34


Kemp, W.M., J.M. Testa, D.J. Conley, D. Gilbert, and J.D. Hagy. 2009. Temporal responses of coastal hypoxia to nutrient loading and physical controls. Biogeosciences 6: 2985-3008.


Testa, J.M., W.M. Kemp, W.R. Boynton, and J.D. Hagy III. 2008. Long-term changes in water quality and productivity in the Patuxent River estuary: 1985 to 2003. Estuaries and Coasts 31: 1021-1037.

Ball, W.P., D.C. Brady, M.T. Brooks, R. Burns, B.E. Cuker, D.M. Di Toro, T.F. Gross, W.M. Kemp, L. Murray, R.R. Murphy, E. Perlman, M. Piasecki, J.M. Testa, I. Zaslavsky. 2008. A Prototype System for Multi-Disciplinary Shared Cyberinfrastructure – Chesapeake Bay Environmental Observatory (CBEO). Journal of Hydrologic Engineering 13(10): 960-970.

Testa, J.M. and W.M. Kemp. 2008. Regional, seasonal, and inter-annual variability of 
biogeochemical processes and physical transport in a partially stratified estuary: a box-modeling analysis. Marine Ecology Progress Series 356: 63-79.


Newell R.I.E., W.M. Kemp, J.D. Hagy, C.F. Cerco, J.M. Testa, and W.R. Boynton. 2007. Top-down control of phytoplankton by oysters in Chesapeake Bay, USA: Comment on Pomeroy et al. (2006). Marine Ecology Progress Series 341: 293-298.


Testa, J.M., M.A. Charette, E.R. Sholkovitz, M.C. Allen, A. Rago, and C.W. Herbold. 2002. Cycling of dissolved iron in the subterranean estuary of a coastal bay: Waquoit Bay, Massachusetts. Biological Bulletin 203(2): 255-256.